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The members of this genus of plant viruses are similar to the Subterranean clover stunt virus. They have a multipartite genome with 6-8 circular single-stranded DNAs each of about 1 kb in size. The ssDNAs are individually encapsidated in small icosahedral virions (18-20 nm) that are transmitted by certain aphid species in a circulative but non-propagative manner. Each DNA carries only a single gene. They are transcribed unidirectionally and contain a replication origin, part of which is a short inverted repeat sequence flanking a conserved sequence of 9 nucleotides.
Currently, the following species have been entered into the system: