Puccinia jaceae (weed pathogen)

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symptoms of Puccinia jaceae var. solstitialis (click on image to enlarge it)
Author(s): Stephen Ausmus, USDA
Source: Wikimedia Commons
symptoms of Puccinia jaceae var. solstitialis on Centaurea solstitialis (click on image to enlarge it)
Author(s): Eric Coombs, Oregon Department of Agriculture
Source: IPM Images

Puccinia jaceae (weed pathogen) G.H. Otth 1865

This rust fungus is a biocontrol agent against Centaurea spp..

Puccinia jaceae var. diffusa - a pathogen of diffuse knapweed (Centaurea diffusa)
Puccinia jaceae var. solstitialis - a pathogen of yellow starthistle (Centaurea solstitialis)

An isolate of Puccinia jaceae var. solstitialis originating from Turkey has been released in western U.S.A. since 2003 and has become established. The fungus is characterized by two supraequatorial germpores on the urediniospores. These spores are golden-brown and slightly elliptical, about 25 x 24 µm in size.

For details see the respective page in Wikipedia.